But there is no space for Manu Tuilagi in the Red Rose 23 – with prop Ben Moon starts his first ever Test in only his second Red Rose cap.
Sam Underhill comes in for injured Tom Curry in the back row in the third and final change to Eddie Jones' side.
Jones knows he has to score tries to crush New Zealand at Twickenham and Sale Sharks star Ashton has 19 in 40 caps for his country.
The 31-year-old comes straight into the side with Exeter Chiefs and Lions winger Jack Nowell dropping to the bench with no room for Tuilagi.
Tuilagi missed out in the win against South Africa after tweaking his groin despite originally being named on the bench to face the Springboks.
Courtney Lawes has replaced Zach Mercer on the bench as Jones beefs up his replacements.
The rest of the subs remains the same with Alec Hepburn dropping down after Moon's promotion to the starting XV.
15 Elliot Daly (Wasps, 22 caps)
14 Chris Ashton (Sale Sharks, 40 caps)
13 Henry Slade (Exeter Chiefs, 14 caps)
12 Ben Te’o (Worcester Warriors, 14 caps)
11 Jonny May (Leicester Tigers, 38 caps)
10 Owen Farrell (Saracens, 62 caps) co-captain
9 Ben Youngs (Leicester Tigers, 78 caps)
1 Ben Moon (Exeter Chiefs, 1 cap)
2 Dylan Hartley (Northampton Saints, 94 caps) co-captain
3 Kyle Sinckler (Harlequins, 14 caps)
4 Maro Itoje (Saracens, 23 caps)
5 George Kruis (Saracens, 26 caps)
6 Brad Shields (Wasps, 3 caps)
7 Sam Underhill (Bath Rugby, 6 caps)
8 Mark Wilson (Newcastle Falcons, 5 caps)
16 Jamie George (Saracens, 29 caps)
17 Alec Hepburn (Exeter Chiefs, 3 caps)
18 Harry Williams (Exeter Chiefs, 12 caps)
19 Charlie Ewels (Bath Rugby, 7 caps)
20 Courtney Lawes (Northampton Saints, 65 caps)
21 Danny Care (Harlequins, 82 caps)
22 George Ford (Leicester Tigers, 48 caps)
23 Jack Nowell (Exeter Chiefs, 27 caps)
Jones said: “The expectation for Saturday is no different to any other Test match. We want to be at our best, better than we were in the previous Test match and we want to play with pride and passion which ignites the fan.
"It’s been a good week, the players have recovered well, trained well on Tuesday, exceptionally well on Wednesday and we look forward to the challenge of taking on New Zealand on Saturday.
“The crowd last week was wonderful, we had great support and enthusiasm for the game and they saw a great Test match and I’m sure on Saturday they will see another.”
1. Karl Tu'inukuafe (10)
2. Codie Taylor (39)
3. Owen Franks (104)
4. Samuel Whitelock (106)
5. Brodie Retallick (72)
6. Liam Squire (21)
7. Ardie Savea (32)
8. Kieran Read – captain (115)
9. Aaron Smith (80)
10. Beauden Barrett (70)
11. Rieko Ioane (21)
12. Sonny Bill Williams (50)
13. Jack Goodhue (5)
14. Ben Smith (74)
15. Damian McKenzie (20)
16. Dane Coles (57)
17. Ofa Tuungafasi (23)
18. Nepo Laulala (14)
19. Scott Barrett (26)
20. Matt Todd (15)
21. TJ Perenara (52)
22. Richie Mo'unga (5)
23. Ryan Crotty (42)
All Blacks boss Steve Hansen said: "There's huge anticipation at what lies ahead on Saturday and the team is certainly looking forward to the challenge of the weekend.
"Our week's preparation has been very good. We've spent the early part of our week getting our preparation right and clarity of our roles and been building throughout the week so we're ready to go on Saturday.
"We know we'll be up against a very determined and well-coached side. They'll have gained a lot of confidence from their win over South Africa and be keen to make a statement in front of their passionate Twickenham home crowd.
"However, for us it's about making our own statement. To do that, we'll need to bring our own intensity and accuracy to an even higher level throughout the game.
"It's a great opportunity to assess where our game is at and we're really looking forward to it."
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